Гербарий БИН РАН - Образцы

Orchidaceae

Dendrobium nobile Lindl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Clustering epiphyte on tall mossy trees. Flowers white, tepal tips purple, lip white with purple violet center. Occasional.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4252 (NHC057).

Orchidaceae

Pholidota chinensis Lindl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Clustering epiphyte on tall mossy trees. Locally very common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4251 (NHC056).

Orchidaceae

Acanthophippium striatum Lindl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial herb on shady slope along forest trail. Flowers white, sepals with purple veins. Locally common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4250 (NHC055).

Convallariaceae

Tupistra muricata (Gagnep.) N.Tanaka?

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial rosulate herb on shady slope along forest trail. Young fruits bright grassy green. Locally common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4249 (NHC054).

Annonaceae

Miliusa balansae Finet & Gagnep.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Treelet about 5-6 m tall on shady slope. Young fruits red. Not common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4248 (NHC053).

Apocynaceae

Rauvolfia verticillata (Lour.) Baill.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Treelet about 2 m tall on shady slope. Flowers white. Not common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4247 (NHC052).

Euphorbiaceae

Antidesma fordii Hemsl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Tree to 10 m tall on shady slope. Not rare.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4246 (NHC051).

Polypodiaceae s.l.

Pyrrosia adnascens (Sw.) Ching

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Creeping epiphyte on tall mossy trees. Very common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4245 (NHC050).

Acanthaceae

Hemigraphis sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial herb in open place along forest trail. Occasional. Spicy plant in local kitchen.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4244 (NHC049).

Polypodiaceae s.l.

Colysis digitata Ching?

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial and lithophytic creeping fern on shady slope. Not rare.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4243 (NHC048).

Icacinaceae

Gomphandra tetrandra (Wall.) Sleumer

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Shrub to 2 m tall on shady slope. Flowers white. Locally common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4242 (NHC047).

Convallariaceae

Ophiopogon latifolius L.Rodr.?

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial herb on shady slope with erect stem to 25 cm long supported by several rigid stilt roots. Flowers white. Locally common.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4241 (NHC046).

Begoniaceae

Begonia lancangensis S.H.Huang

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb to 1 m tall on shady slope along trail. Flowers white. Leaves with dark purple nerves. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4224 (NHC035).

Zingiberaceae

Amomum sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial herb with leaves to 1.2 m long in open place along trail. Flowers bright yellow. Occasional.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4234 (NHC045).

Araceae

Raphidophora decursiva (Roxb.) A. Schott

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Creeping epiphytic semi-woody vine to 10 m long on tall trees. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4233 (NHC044).

Annonaceae

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 6-7 m tall on shady slope. Flowers dark green. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4232 (NHC043).

Theaceae

Gordonia axillaris (Roxb.) D.Dietr.?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 10 m tall on shady slope. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4231 (NHC042).

Lauraceae

Litsea baviensis Lecomte

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 10 m tall on shady slope. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4230 (NHC041).

Myrsinaceae

Ardisia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Shrub to 2 m tall on shady slope. Flower buds dull pinkish. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4229 (NHC040).

Urticaceae

Elatostema xanthophyllum W.T.Wang?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial herb to 0.4 m tall on shady slope. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4228 (NHC039).

Rubiaceae

Mussaenda sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Scandent shrub of vine to 3 m long on open steep slope along trail. Floral bracts white. Very common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4227 (NHC038).

Saurauiaceae

Sauraia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Tree about 15 m tall on shady slope along trail. Flowers light pink. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4226 (NHC037).

Moraceae

Ficus sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Woody vine on shady slope along trail. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4225 (NHC036).

Begoniaceae

Begonia lancangensis S.H.Huang

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb to 1 m tall on shady slope along trail. Flowers white. Leaves with dark purple nerves. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4224 (NHC035).

Commelinaceae

Pollia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb to 0.8 m tall in open wet place. Very common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4223 (NHC034).

Apocynaceae

Tabernaemontana sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet about 3.5 m tall on steep slope. Fruits orange. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4222 (NHC033).

Myrsinaceae

Ardisia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet about 3.5 m tall on steep slope. Flower buds light purple. Not rare.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4221 (NHC032).

Rubiaceae

Myrioneuron effusum (Pit.) Merr.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Undershrub 2-2.5 m tall on steep slope. Flowers yellow. Not rare.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4220 (NHC031).

Symplocaceae

Symplocos sp.?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 4 m tall. Flowers yellow. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4219 (NHC030).

Annonaceae

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet or shrub 2-3 m tall. Flowers yellow, fruits green. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4218 (NHC029).

Moraceae

Ficus sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Tree to 8 m tall in shady slope. Syconia green. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4217 (NHC028).

Rubiaceae

Psychotria sp.?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Shrub or treelet to 3-4 m tall. Flowers white. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4216 (NHC027).

Rubiaceae

Tarenna sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Shrub or treelet to 3-4 m tall. Flowers white to greenish. Fruits green. Occasional.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4215 (NHC026).

Euphorbiaceae

Claoxylon indicum (Blume) Hassk.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 5-6 m tall. Flowers white. Occasional.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4214 (NHC025).

Berberidaceae

Dysosma versipellis (Hance) T.S.Ying

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial rhizomatous herb in shady places. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4213 (NHC024).

Orchidaceae

Phalaenopsis chibae T.Yukawa

The border on Kon Tum and Quang Nam Provinces along the road Da Nang – Dak Gley. Secondary broad-leaved evergreen forest on very steep slopes composed by silicate shale at elevation of 800–1100 m a.s.l. Epiphyte on mossy lianas on shady place on steep stream slope. Rare.

03 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4142.

Flowered in cultivation on 13 June 2025, Nguyen Van Canh, AL4142a.

Taccaceae

Tacca chantrieri André

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial rosulate herb in shady places. Floral bracts green with violet, flowers light green. Common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4212 (NHC023).

Convallariaceae

Polygonatum punctatum Kunth

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial herb to 0.5 m tall on steep rocky slope. Rare.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4211 (NHC022).

Asclepiadaceae

Hoya multiflora Blume

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Epiphytic undershrub with pendulous and erect shoots to 1.5 m long on tall mossy tree. Flowers buds white. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4210 (NHC021).

Meliaceae

Aglaia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Tree to 20 m tall and 40 cm DBH. Fruits globular to 3 cm in diameter, dull gray brownish. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4209 (NHC020).

Euphorbiaceae

Phyllanthus sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 6 m tall. Occasional.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4208 (NHC019).

Phormiaceae

Chlorophytum malayense Ridl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial rosulate herb in open wet place along trail. Flowers white, anthers yellow. Occasional.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4207 (NHC018).

Polypodiaceae

Leptochilus sp.?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Lithophytic creeping fern in shady place. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4206 (NHC017).

Gesneriaceae

Rhynchotechum ellipticum (D.Dietr.) A.DC.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb or undershrub to 1.5 m tall in open place along trail. Not rare.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4205 (NHC016).

Asclepiadaceae

Dischidia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Herbaceous creeping epiphytic wine to 1.5 m long on tall tree. Flowers white, Common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4204 (NHC015).

Verbenaceae

Gomphostemma leptodon Dunn?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb or undershrub to 2 m tall in open place along trail. Flowers white. Common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4203 (NHC014).

Araceae

Aglaonema sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Erect terrestrial herb to 1 m tall in open place along trail. Spathe light green, inflorescence white. Very common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4202 (NHC013).

Selaginellaceae

Selaginella sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial herb to 0.6 m tall in open place along trail. Not common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4201 (NHC012).

Rubiaceae

Ophiorrhiza sanguinea Blume?

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Terrestrial herb to 0.5 m tall in open wet place. Flowers white to pinkish. Not common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4200 (NHC011).

Myrsinaceae

Ardisia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Undershrub to 1 m tall in open place along road. Flowers pink purple. Rare.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4199 (NHC010).