Гербарий БИН РАН - Образцы

Orchidaceae

Diploprora carinata Aver. & V.C.Nguyen, sp. nov.

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1692.

Plant originated from Quang Nam Province, Nong Son District, evergreen broad-leaved forest at elevation 100–150 m a.s.l. 2022, Nguyen Van Canh.

Plant flowered in culture on 23 March 2026, L. Averyanov, AL1692.

All photos © L. Averyanov

Orchidaceae

Diploprora carinata Aver. & V.C.Nguyen, sp. nov.

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1692.

Plant originated from Quang Nam Province, Nong Son District, evergreen broad-leaved forest at elevation 100–150 m a.s.l. 2022, Nguyen Van Canh.

Plant flowered in culture on 23 March 2026, L. Averyanov, AL1692.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus

Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, to the EEN of Bai Dinh village around point 17°45’27”N 105°47’46”E. Primary broad-leaved closed wet forest on very steep rocky slopes and cliffs of low remnant mountains composed with highly eroded solid marble-like limestone at elevation about 600-700 m a.s.l. Tree about 6 m tall near mountain top. Rare.

HAL 12507 11 February 2009 Photo.

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Vietnam, Thua Thien – Hue Prov., Huong Thuy Distr., Duong Hoa Municipality, Huong Thuy forest enterprise territory, around point 16°13'20''N, 107°34'23''E, SW slopes of Mang Chan ridge at elevation 600-800 m a.s.l. Fractionally logged primary closed evergreen broad-leaved forest along tops of ridge composed with rocky quartzite outcrops. Tree 25-35 m tall. Occasional emergent of canopy forest stratum.

No HAL 8054 10 May 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, N.T.Vinh.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Vietnam, Thua Thien – Hue Prov., A Luoi Distr., A Roang Municipality, Tra Lenh Forestry Department station, around point 16°04'38''N, 107°29'10''E. Primary closed evergreen broad-leaved forest on very steep mountain slopes composed with shale and granite at elevation 700-800 m a.s.l. Seedlings of tree up to 30 m tall and 1.2 m in diam. Occasional co-dominant of first forest stratum along tops of ridge.

No HAL 7179 20 April 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, J.Regalado, T.V.Thao, N.T.Vinh.

All photos © L. Averyanov

Podocarpaceae

Dacrydium elatum

S. Vietnam, Gialai, Kbang,

1989, L. Averyanov.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus

S. Vietnam, Gialai, Kbang,

1989, L. Averyanov.

All photos © L. Averyanov

FLORA OF VIETNAM

Fam. Pinaceae

Keteleeria sp.

N. Vietnam, prov. Cao Bang, distr. Tra Linh, municipality Luu Ngoc, vicinities of Lung Mu village (N 22ø47' E 106ø18'), about 14 km from Cao Bang town to the N. Primary coniferous and mixed semideciduous open forest (with Pseudotsuga brevifolia) along tops of limestone ridge at 800 m alt. Tree up to 20-25 m hg. Main forest codominante (with age up to 600-800 eyars).

No CBL 1196 13 Dec. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, N.T.Hiep, P.K.Loc, N.X.Tam.

All photos © L. Averyanov

Plants of Ta Dung Nature Reserve

Podocarpaceae

Dacrycarous imbricatus

Vietnam, Dak Nong Prov., Dak Glong Distr., Dak Plao Municipality, steep mountain slopes of Ta Dung mountain system at elevation 1000-1250 m a.s.l. around point 11º52’30’’N 108º01’12’’E. Primary broad-leaved evergreen closed wet forest on SW slopes composed with crystalline highly metamorphosed sandstone. Sapling 1 m tall of tree up to 25 m high. Locally common.

HLF 5556 9 November 2005. Photo.

Coll. L.Averyanov, P.K.Loc, T.V.Thao, N.T.Vinh.

Grant from Henry Luce Foundation.

All photos © L. Averyanov

Plants of Chu Yang Sin National Park

Podocarpaceae

Dacrycarpus inbricatus

Vietnam, Dak Lak Prov., Krong Bong Distr., Hoa Son Municipality, E slopes of Chu Yang Sin mt system at elevation 600-850 m a.s.l. around point 12º30’27’’N 108º22’50’’E. Secondary broad-leaved evergreen forest with bamboo on steep rocky mountain slopes. Tree to 30 m tall.

HLF 5486a 30 October 2005. Photo.

Coll. L.Averyanov, T.V.Thao, N.T.Vinh.

Grant from Henry Luce Foundation.

All photos © L. Averyanov

Plants of Chu Yang Sin National Park

Podocarpaceae

Dacrycarpus imbricatus

Vietnam, Dak Lak Prov., Krong Bong Distr., Hoa Son Municipality, vicinities of Krong Kmar river, NE slopes of Chu Yang Sin mt system at elevation about 800-1000 m a.s.l. Primary mixed broad-leaved and coniferous forest with Pinus krempfii, Dacrycarpus imbricatus and Dacrydium elatum along ridge edge. Treelet 3-4 m tall of tree up to 20-25 m tall. Common. Co-dominant of first forest stratum.

HLF 5420 28 October 2005. Photo.

Coll. L.Averyanov, T.V.Thao, N.T.Vinh.

Grant from Henry Luce Foundation.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°38’00.4”N 105°55’57.9”E, Kalap Mountain. Primary mixed and coniferous forest (with Dacrydium elatum and Dacrycarpus imbricatus) on very steep rocky slope and on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation 750-800 m a.s.l. Tree to 35 m tall, to 60 cm DBH on steep shady rocky slope near to mountain top and on top. Co-dominant of coniferous forest.

CPC 3726 25 July 2011

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

N.Vietnam, Thanh Hoa Prov., Ba Thuoc Distr., Co Lung Municipality, Khuyn village, around point 20º26'31''N, 105º14'55''E. Primary evergreen seasonal broad-leaved submontane forest on crystalline marble-like highly eroded limestone, slopes of limestone ridge near the top at elev. 700-800 m. Tree 40-45 (or more) m tall. Occasional emergent of the 1 forest stratum.

No HAL 3078 18 Sept. 2003 (Photo)

FFI Pu Luong - Cuc Phuong Conservation Project

Collectors: L.Averyanov, D.T.Doan, J.Regalado, N.T.Vinh.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus

Son La Prov., Thuan Chau Distr., Co Ma Municipality, Hua Ty village, Copia nature reserve, around point 21º20’07’’N 103º35’07’’E. Remnants of broad-leaved (former coniferous with Fokienia hodginsii) primary and secondary forest on steep slopes and on tops of ridge composed with clay shale at elev. 1500-1600 m a.s.l. Tree up to 35 m tall and 1.2 m in diam. Not common. Former co-dominant of coniferous forest.

HAL 9713 12 Nov. 2006 DNI fix.

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus

Lai Chau Prov., Sin Ho Distr., Ma Quai Municipality, Can Ti 2 village, around point 22º24’18’’N 103º22’45’’E. Remnants of broad-leaved evergreen closed primary forest on steep mountain slopes composed with sandstone at elev. about 1100 m a.s.l. Tree 10-12 m tall. Rare.

HAL 9859 17 Nov. 2006 Photo.

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Saravan Prov., Sa Mouay Distr., to the SEE of Sa Mouay town, elevation from Ta Oy village (700 m elev.) to mountain ridge with highest point at 1350 m around 16°08’47.0”N 106°52’35.4”E. Primary evergreen humid mixed and coniferous forest on slopes composed with silicate rocks. Tree to 40 m tall and to 1 m DBH on tops and on ridge slopes from 1150 to the tops of ridge. Common. Significant co-dominant of coniferous forest.

LA-VN 197 04 March 2013

Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

All photos © L. Averyanov

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Kham Mouan Prov., Bounlapha Distr., Thong Sam village, Hinnam No Protected Area, Pu Pha Song Mt. composed by sandstone, around point 17°35’13.2”N 105°47’49.8”E. Primary and secondary evergreen broad-leaved and mixed forest on W macroslope at elevation from 500 m to the mountain top elevated to 1425 m. Tree 20-30 m tall to 0.6 m DBH. Very common, forest co-dominant at elevations 700-1000 m.

LA-VN 238 09 March 2013

Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

All photos © - L. Averyanov

Pinaceae

Tsuga chinensis

Tuyen Quang, Na Hang, Sinh Long, P.V.The, 2012.

All photos © The Van Pham

Pinaceae

Tsuga chinensis (Franch.) Pritz.

Northern Vietnam, Ha Giang Prov., Quan Ba Distr., Thai An Municipality, in the vicinity of Lo Thang village, around point 22ø59'50''N, 105ø05'46''E. Primary coniferous forest with Tsuga chinensis on remnant highly eroded limestone ridge at elev. about 1600 m a.s.l. Tree 15-20 m hg up to 0.5 m in diam. Co-dominant of coniferous forest on tops of ridges.

No HAL 1503 (Photo) 10 May 2002

Collectors: P.K.Loc, L.Averyanov, N.T.Vinh.

All photos © L. Averyanov

Pinaceae

Tsuga chinensis

Vietnam, Ha Giang Prov., Dong Van Distr, Ta Lung Municipality, vicinities of Xa Lung village around point 23º14’50’’N, 105º21’34’’E at elevation about 1450-1550 m a.s.l. Degraded primary coniferous forest with Pseudotsuga sinensis, Tsuga chinensis and Pinus sp. on very steep rocky slopes and along tops of narrow rocky remnant ridge composed with highly eroded marble-like limestone. Tree up to 15 m tall on very steep rocky mossy slope. Co-dominant of coniferous forest.

No HAL 8589 14 Dec 2005. Photo.

Collectors: L.Averyanov, T.V.Thao, P.V.The.

All photos © - L. Averyanov

Pinaceae

Tsuga chinensis

N. Vietnam, prov. Ha Giang, distr. Yen Minh, municipality Lao Va Chai, vicinities of Ngan Chai village, 6 km to W of Yen Minh town (N 23ø07' E 105ø08'). Primary evergreen wet mossy mixed forest (with domination of Tsuga chinensis and Pinus kwangtungensis) along tops of karst remnant limestone ridge at 1500-1600 m elev. Tree 15-20 m hg. Very common. Main codominante of 1 forest stratum.

No CBL 1885 01 May 1999

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: P.K.Loc, P.H.Hoang, Averyanov L.

All photos © L. Averyanov

Pinaceae

Tsuga chinensis (Franch.) Pritz.

Northern Vietnam, Ha Giang Prov., Quan Ba Distr., Can Ty Municipality, in the vicinity of Lung Vai village at left side of Mien River, around point 23ø05'09''N, 105ø03'23''E. Primary coniferous forest with Pseudotsuga brevifolia and Tsuga chinensis on remnant highly eroded limestone ridge at elev. 1440 m a.s.l. Tree 15-20 m hg and up to 40 cm in diam. Co-dominant of first forest stratum.

No HAL 1448 (Photo) 8 May 2002

Collectors: P.K.Loc, L.Averyanov, N.T.Vinh.

All photos © - L. Averyanov

Pinaceae

Tsuga chinensis (Franch.) Pritz.

Northern Vietnam, Ha Giang Prov., Quan Ba Distr., Can Ty Municipality, in the vicinity of Lung Vai village at left side of Mien River, around point 23ø05'09''N, 105ø03'23''E. Primary coniferous forest with Pseudotsuga brevifolia and Tsuga chinensis on remnant highly eroded limestone ridge at elev. 1440 m a.s.l. Tree 15-20 m hg and up to 40 cm in diam. Co-dominant of first forest stratum.

No HAL 1448 (Photo) 8 May 2002

Collectors: P.K.Loc, L.Averyanov, N.T.Vinh.

All photos © - L. Averyanov

Pinaceae

Tsuga dumosa

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Yen Lac, vicinities of Yen Lac village (N 22ø44' E 105ø50'), about 44 km to NWW from Cao Bang town. Primary open coniferous mossy forest along tops of

limestone remnant karst ridge at 1500 m alt. Tree 15-20 m hg. Codominante of 1 forest stratum.

No CBL 246 15 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., N.T.Hiep, P.K.Loc, N.X.Tam.

All photos © L. Averyanov

Pinaceae

Tsuga chinensis (Franch.) Pritz.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary mixed and coniferous humid evergreen forest (with Taxus, Podocarpus, Tsuga, Pseudotsuga and Xanthocyparis) on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1200-1250 m a.s.l. around point 22°50’03.8’’N 105°51’36.5’’E. Tree 4-5 m tall and 10 cm DBH on ridge tops. Not very common.

20 November 2014 CPC 7541

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

All photos © L. Averyanov

Orchidaceae

Hetaeria nitida Ridl.

Gia Lai Province, K'Bang District, natural forest, alt.: 670 m nearby waterfall.

22 March 2026, Dinh Quang Diep, AL4545.

All photos © Dinh Quang Diep

Orchidaceae

Zeuxine parvifolia (Ridl.) K.Schum. & Fedde

Da Nang, Son Tra,

2 March 2026, L. Averyanov, AL4544.

All photos © Nguyen Van Canh

Orchidaceae

Zeuxine affinis Hook.f.

Da Nang, Son Tra,

2 March 2026, L. Averyanov, AL4543.

All photos © Nguyen Van Canh

Orchidaceae

Cheirostylis yunnanensis Rolfe

Da Nang, Son Tra,

2 March 2026, L. Averyanov, AL4542.

All photos © Nguyen Van Canh

Pinaceae

Tsuga chinensis (Franch.) Pritz.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1500-1550 m a.s.l. around point 22°50’01.3’’N 105°50’05.7’’E. Tree 20-30 m tall along mountain tops. Very common. Usual dominant of mixed and coniferous forest. Widewly logged.

20 November 2014 CPC 7508 photos

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

All photos © - L. Averyanov

Pinaceae

Tsuga canadensis

Ccult. USA Washington area

Feb. 2008, L. Averyanov.

All photos © - L. Averyanov

Pinaceae

Tsuga canadensis

St. Louis, Missouri Botanical Garden, U.S.A.

2003, L. Averyanov.

All photos © - L. Averyanov

Pinaceae

Pseudotsuga sinensis

Vietnam, Ha Giang Prov., Meo Vac Distr, Pai Lung Municipality, around point 23º15’06’’N, 105º22’02’’E at elevation about 1300 m a.s.l. Highly degraded remnants of coniferous forest with Pseudotsuga sinensis and Tsuga chinensis on very steep rocky slopes and along tops of rocky remnant ridges composed with highly eroded marble-like limestone. Tree up to 15 m tall. Co-dominant of coniferous forest.

No HAL 8547 12 Dec 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, P.V.The, N.S.Khang.

All photos © - L. Averyanov

Pinaceae

Pseudotsuga sinensis Dode var. brevifolia (C.Y.Cheng, W.C.Cheng et L.K.Fu) Farjon et J.Silba

(Pseudotsuga brevifolia C.Y.Cheng, W.C.Cheng et L.K.Fu)

Vietnam, Ha Giang Prov., Meo Vac Distr, Sung Tra Municipality, vicinities of Ta Cha Lang village around point 23º10’12’’N, 105º17’39’’E at elevation about 1100-1150 m a.s.l. Highly degraded remnants of coniferous forest with Tsuga chinensis on very steep rocky slopes and along tops of rocky remnant ridges composed with highly eroded marble-like limestone. Young treelet 3-4 m tall. Not rare.

No HAL 8506 11 Dec 2005.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, P.V.The, N.S.Khang.

All photos © - L. Averyanov

Pinaceae

Pseudotsuga brevifolia

Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Nhieu Lung village, around point 22°46'07.9"N, 105°53'59.4"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation 1150-1250 m a.s.l. Shrub 203 m tall on rather open rocky slope. Flowers white with light purple tint. Very common, particularly among open secondary scrub.

8 October 2013 CPC 5476a

L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

All photos © - L. Averyanov

Pinaceae

Pseudotsuga sinensis Dode

Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Ngan Vai village, around point 22°46'53"N, 105°52'58.7"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1300 m a.s.l. Tree to 15 m tall along mountain ridge and on high part of mountain slope. Very common, main dominant of the forest.

7 October 2013 CPC 5436

L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

All photos © - L. Averyanov

Pinaceae

Pseudotsuga sinensis Dode

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village. Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45’24.8’’N 105°46’47.9’’E. Tree to 7 m tall and 10 cm DBH on mountain tops. Very common. Usual co-dominant of coniferous forest.

24 November 2014 CPC 7635

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

All photos © - L. Averyanov

Pinaceae

Pseudotsuga brevifolia C.Y.Cheng, W.C.Cheng & L.K.Fu

Northern Vietnam, Ha Giang Prov., Quan Ba Distr., Can Ty Municipality, in the vicinity of Sin Suoi Ho village at right side of Mien River, about 1 km to the N from point 23ø07'12''N, 105ø00'52''E. Primary coniferous forest with Pseudotsuga brevifolia on remnant highly eroded limestone ridge at elev. 1000-1200 m a.s.l. Tree 10-20 m hg. Monodominant of first forest stratum along tops of ridges. Extensively logging.

No HAL 1548 (Photo) 11 May 2002

Collectors: P.K.Loc, L.Averyanov, N.T.Vinh.

All photos © - L. Averyanov

Pinaceae

Pseudotsuga brevifolia

N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 800-850 m a.s.l. Remnants of primary broad-leaved and coniferous forest on ridge tops and on very steep slopes and cliffs of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Tree 6-10 m tall. Not common due to total logging. Former co-dominant of primary closed coniferous forest that extinct here 15-20 years ago.

No HAL 5499 9 June 2004 (Photo)

Collectors: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

All photos © - L. Averyanov

Pinaceae

Pseudotsuga brevifolia

N.Vietnam, Bac Kan Prov., Na Ri Distr., Liem Thuy Municipality, village Na Bo, around point 21º56'44''N, 106º05'09''E, at elevation 300-700 m a.s.l. Primary broad-leaved evergreen closed wet forest on very steep slopes and cliffs of rocky ridges and mesae composed with white solid marble-like crystalline limestone. Tree 8-15 m tall on tops of ridge. Common. Occasional co-dominant of fragments of coniferous forest on tops of ridges.

No HAL 4917 27 May 2004

Collectors: L.Averyanov, N.T.Hiep, P.V.The, N.T.Vinh.

All photos © - L. Averyanov