Гербарий БИН РАН - Образцы

FLORA OF LAOS

Polypodiaceae s.l.

Pseudodrynaria sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Short creeping epiphyte on tall trees. Very common.

LA-VN 1228 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Belvisia sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.

LA-VN 1339 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium rupestre Hope

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial and epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.

LA-VN 1338 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Orchidaceae

Tainia penangiana Hook.f.

Dak Lak Province, Lak District, around Lieng Bok Sak Waterfall at elevation about 400 m a.s.l. Dry evergreen broad-leaved forest along rocky stream on granite. Terrestrial herb among rocks along stream. Occasional.

11 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3144.

Flowered in culture on 12 January 2026, L. Averyanov, T. Maisak, AL3144.

Acanthaceae

Gen. ?

Dak Lak Province, M’Drak District, W foothills of Chu Mu Mt., dry secondary and remnants of primary evergreen broad-leaved forest on granite on steep slopes along rocky river at elevation 600-650 m a.s.l. Terrestrial adpressed rosulate herb on soil cliff in shady place along stream. Flowers light blue violet. Not rare.

25 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL3944.

Convallariaceae

Peliosanthes sp.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l.

Terrestrial herb in shady rocky place. Locally common.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2385.

VIETNAM

POLYPODIACEAE

Pyrrosia

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

Epiphytic, vine. Fronds light green;

sporangia orange-brown; abaxial leaf

surfaces very light brown.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6652

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

POLYPODIACEAE

Lepisorus

Lao Cai

Van Ban District; Nam Xe Municipality;

Nam Xi Tan Village. Collection in forest

on steep W-facing slope along stream;

just above village; on N side of road

between Nam Xe and Khau Co Pass at near

40 km to Than Uyen mark.

22°02'31"N 103°58'32"E 700 - 1113 m

Epiphytic, on slope in shade,

occasional, creeper. Rhizome green with

26 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz, S. Bodine & N.Q. Hieu 6944

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

brown hairs; sporangia light

yellow-green.

VIETNAM

POLYPODIACEAE

Lepisorus subrostratus (C. Chr.) C. Chr.

& Tardieu

det. P.K. Loc, 2001

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

Occasional, epiphytic. Pinnae light

green; sporangia light brown.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6650

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

POLYPODIACEAE

Colysis

Son La

Yen Chau District; Muong Lum

Municipality; Lum Village. Collections

on ridge S of village. Collected with

P.K. Loc, N.T. Hiep, L.V. Averyanov,

G.E. Schatz, S. Bodine & N.Q. Hieu.

21°00'47"N 104°29'30"E 900 - 1134 m

Occasional, lithophytic. Rhizome green;

rachis green; sporangia dark brown.

2 March 2001

D.K. Harder & et al. 7115

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

POLYPODIACEAE

Colysis

Son La

Yen Chau District; Muong Lum

Municipality; Lum Village. Collection on

ridges S of village. Collected with P.K.

Loc, N.T. Hiep, B. Tan, G.E. Schatz,

L.V. Averyanov & S. Bodine.

21°00'17"N 104°29'08"E 1242 m

Occasional, lithophytic, on mossy rocks.

Rachis black; sporangia dark dull

black-brown; rhizome green with brown

hairs.

3 March 2001

D.K. Harder & et al. 7215

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

POLYPODIACEAE

Colysis

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000); valley just W of

Bat Dai Son border station; valley of

slash and burn agriculture, then higher

up, degraded secondary vegetation in

shade.

23°09'04"N 104°59'53"E 1000 m

Occasional, among rocks. Hair on rhizome

11 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6146

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

brown; sporangia dark brown.

VIETNAM

POLYPODIACEAE

Colysis

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophyte herb in shady wet places.

Common.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7412

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae s.l.

Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1500-1700 m. Creeping epiphyte at the base of mossy trees. Very common.

No VH 6040 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,

Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1300 m. Creeping epiphyte at base of large trees. Very common.

No VH 6240 08.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Plagyogeria sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1700-1750 m. Terrestrial fern on very steep shady slopes of river

canyon. Not rare.

No VH 6368 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Plagyogeria sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,

Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1550 m. Terrestrial fern on shady very steep slope of river

canyon. Not rare.

No VH 6248 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

ASPLENIACEAE

Asplenium

Ha Giang

Vi Xuyen District; Minh Tan Municipality

Ban Ngay Village. Collection on E

facing ridge above river; on exposed

limestone rocks; collections in

disturbed agricultural land leading into

primary forest with some deforestation

of Burretiodendron (Tiliaceae) trees.

23°00'09"N 104°55'21"E 225 - 621 m

23°00'37"N 104°54'46"E

Frequent epiphyte and occasional

17 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz & S. Bodine 6464

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

lithophyte. Fronds light green;

sporangia dull, light brown; midrib of

frond purple-brown, producing "nests"

when large.

VIETNAM

ASPLENIACEAE

Asplenium

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophytic fern. Common.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7392

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Asplenium antrophyoides Christ

Northern Vietnam, Thanh Hoa, Pu Luong Nature Reserve.

2019

Coll. : Cuong Huu Nguyen CLL 529

Det. Averyanov L. 26.12.2025

VIETNAM

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Epiphyte in shady places. Common.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7411

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

evergreen, closed forest on limestone

ridge and cliffs, some coniferous forest

with Pinus kwangtungensis and Calocedrus

macrolepis.

20°46'35"N 104°46'31"E 1400 - 1500 m

6 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7335

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

---------------------------------------

Page 2

Epiphyte in shady place. Not rare.

POLYPODIACEAE

Pyrrosia

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000). On limestone

ridge of degraded forest just northwest

of Bai Dai Son border station;

collections in forest above cultivations

(Forest possibly burned ca. 25 years ago

or so, only small trees and few fire

tolerant herbs).

13°09'13"N 104°59'36"E 1064 m

10 February 2001

D.K. Harder, Hiep, P.K. Loc, L.V.

Averyanov, G.E. Schatz & S.A. Bodin 6092

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

Epiphytic. Fronds light green; sporangia

rusty-brown.

VIETNAM

GRAMMITIDACEAE

Grammitis

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Rare, lithophytic, in shade. Sporangia

brown.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6359

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Epiphyte in shady places. Common.

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7488

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

Grammitideae C. Presl

Son La

Yen Chau District; Muong Lum

Municipality; On Oc Village; collection

along N-S ridge along western side.

Collection along trail from agricultural

field to secondary vegetation to primary

forest vegetation with some trees

removed along ridge top. Limestone

substrate (crystalline). GPS coordinates

at top of ridge.

20°59'06"N 104°28'46"E 1468 m

4 March 2001

D.K. Harder, P.K. Loc & N.T. Huong 7261

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

Epiphytic. Root brown; raised sporangiabrown.

Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Epiphytic, occasional, near top of small

hill. Rhizome and roots dark, dull

brown; sporangia light brown in small

light green cups.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6379

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae

Asplenium sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system to main peak (12ш24'N, 108ш26'E). Primary broad-leaved mountain evergreen closed

forest on granites along river slopes at elev. 1000-1200 m. Epiphyte fern with pendent fronds up to 30 cm lg on mossy trees along shady

waterfall sides. Very common.

No VH 6021 05-06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2100-2300 m. Epiphyte on mossy trees in shady place. Very common.

VH 6215 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 728, 12.10.1990

POLYPODIACEAE

Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 728, 12.10.1990

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 987, 19.10.1990

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1062, 20.10.1990

POLYPODIACEAE

Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 679, 11.10.1990

FLORA OF VIETNAM

Polypodiaceae s.l.

Pteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.

No VH 6291 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Woodwardtia sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.

No VH 6290 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved forest on granites

along river slopes at elev. 500-900 m. Lithophytic fern on shady mossy river rocks. Common.

No VH 6070 04.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1200-1300 m.

Lithophytic fern on wet mossy shady rocks along river. Common.

No VH 6387 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae

Asplenium aff.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1750 m. Epiphyte at base of large trees on open mossy cliff. Common.

No VH 6318 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Dipteris aff.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1200-1250 m. Lithophytic fern in crevices of open mossy cliffs in river canyon. Not common.

No VH 6276 06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh