Гербарий БИН РАН - Образцы

Orchidaceae

Coelogyne ovalis Lindl.

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Epiphyte.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3496.

Flowered in cultivation 04 December 2025, L. Averyanov, T. Maisak, AL3496.

Orchidaceae

Oberonia pachyrachis Rchb.f. ex Hook.f.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4534.

Orchidaceae

Erythrodes hirsuta (Griff.) Ormerod

Lam Dong Province, Di Linh District, Gia Bac Village. Secondary evergreen broad-leaved and mixed forest (with Pinus kesiya) on granite on very steep slopes at elevation 1000-1100 m a.s.l. terrestrial creeping herb in shady stream slope. Not rare.

29 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3454.

Flowered in culture on 05 February 2026, L. Averyanov, T. Maisak, AL3454.

FLORA OF LAOS

Polypodiaceae s.l.

Cyrtomium sp.

Houphan Prov., Hem Distr., Nacoc village, Puphadeng mountain, around point 20°09'52.6''N 103°10'19.4''E.Primary and secondary broad-leaved evergreen open dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1200 m a.s.l. Terrestrial fern on steep rocky slope near mountain top. Very common.

LA-VN 1425 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF VIETNAM

Polypodiaceae s.l.

Polystichum sp.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1400-1450 m a.s.l. around point 22°49’23.4’’N 105°49’54.5’’E. Lithophytic fern on mossy shady rocks on very steep slope. Common.

19 November 2014 CPC 7495

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Colysis sp.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village. Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45’24.8’’N 105°46’47.9’’E. Lithophytic fern on rocky slopes and on mountain top. Very common.

24 November 2014 CPC 7647

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae s.l.

Polystichum sp.

Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Terrestrial and lithophytic fern along shady rocky ridge edge. Common.

13 November 2014 CPC 7414 a

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Polystichum sp.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1400-1450 m a.s.l. around point 2°49’23.4’’N 105°49’54.5’’E. Lithophytic rosulate fern on mossy shady cliff. Occasional.

19 November 2014 CPC 7499

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Polystichum sp.

Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Lithophytic and terrestrial fern among shady rocks on steep slopes and along ridge edge. Very common.

13 November 2014 CPC 7404

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Polystichum sp.

Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Terrestrial and lithophytic fern along shady rocky ridge edge. Common.

13 November 2014 CPC 7414 b

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Lepisorus sp.?

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest (with Podocarpus, Pinus, Fokienia and Tsuga) on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation about 1500-1550 m a.s.l. around point 22°49’15.4’’N 105°49’53’’E. Creeping epiphyte and lithophyte on rocky mossy mountain slope. Common.

21 November 2014 CPC 7571

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Cyrtomium sp.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village. Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45’24.8’’N 105°46’47.9’’E. Terrestrial and lithophytic fern on steep rocky slope near mountain top. Common.

24 November 2014 CPC 7643

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae s.l.

Colysis sp.

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.

Terrestrial fern on steep shady rocky slope. Common.

LA-VN 1487 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

FLORA OF LAOS

Polypodiaceae s.l.

Cyrtomium sp.

Xiangkhouang Prov., Peak Distr., Khoang village, Pa Khao mt., around point 19°33’02.2’N 103°39’56.3’E. Primary and secondary broad-leaved evergreen dry forest on solid rocky highly eroded limestone at elev. 1200-1450 m a.s.l. Terrestrial and lithophytic fern on very steep slopes and cliffs. Not rare.

LA-VN 933 3 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Colysis sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern in shady slope near mountain base. Common.

LA-VN 1342 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

VIETNAM

THELYPTERIDACEAE

Cyclosorus

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

In shady wet place. Rachis shiny

black-purple, rhizome and rachis hairs

dark brown-black; rachis shiny

purple-black.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6662

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae

Cyclosurus sp.

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14ш39'N, 108ш25'E). Primary broad-leaved wet

evergreen forest on sandstone at elev. 1100-1200 m. Terrestrial fern with fronds up to 2 m lg on steep shady wet slope along forest stream.

Very common.

No VH 5392 19.04.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, P.K.Loc, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Cyathaea sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1400 m. Woody fern 2-3 m hg with fronds up to 2-2.5 m lg in shady wet river side slope.

Not rare.

No VH 6384 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Cyathaea sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1400 m. Woody fern 6 m hg with fronds up to 4-5 m lg in shady wet river side slope. Not

rare.

No VH 6385 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Cyclosurus sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2440 m. Creeping epiphyte. Not common.

No VH 6204 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae

Athyrium sp.

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14ш39'N, 108ш25'E). Primary broad-leaved wet

evergreen forest on sandstone at elev. 1100-1200 m. Terrestrial fern in open wet slope. Common.

No VH 5382 18.04.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, P.K.Loc, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

DRYOPTERIDACEAE

Cyrtomium

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000); ridge N of Bat

Dai Son border station.

23°09'11"N 104°59'06"E 1287 m

Rare, in deep shade, on rocks. Hair on

rachis and lower shoots brown; sporangia

dark brown.

11 February 2001

D.K. Harder, N.T. Hiep, L.V. Averyanov &

P.K. Loc 6181

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

THELYPTERIDACEAE

Cyclosorus

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Occasional, in understory in shade.

Rachis brown; sporangia dark brown.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6395

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

DRYOPTERIDACEAE

Cyrtomium

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Occasional, among rocks. Hairs on

rhizome and rachis brown, shiny;

sporangia brown; rachis green, shiny.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6356

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

CYATHEACEAE

Cyathea

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Collection

on ridge and small limestone peak just

east of house of village head; very

disturbed, secondary vegetation on lower

slopes and on limestone rocks up to

ridgetop.

22°53'11"N 104°56'43"E 437 - 645 m

22°53'23"N 104°56'48"E

Occasional. On stem to 2 m; rachis dull

14 February 2001

D.K. Harder, P.K. Loc, G.E. Schatz &

L.V. Averyanov 6298

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

brown; fronds to 2 m long; pinnae dark

green above; sporangia brown.

VIETNAM

POLYPODIACEAE

Crypsinus

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Epiphyte fern on mossy trees. Common.

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7489

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

THELYPTERIDACEAE

Cyclosorus

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Occasional, in shade. Rachis green,

shiny; sporangia brown.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6607

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

THELYPTERIDACEAE

Cyclosorus

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Occasional, in shade. Pinnae dull dark

green; rachis shiny green to darker

brown at base; sporangia brown.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6624

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

Pteridophyta J.Y. Bergen & B.M. Davis

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Terrestrial and lithophytic fern. Not

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7499

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)